Multicellular chemotrophs?

What characteristics might define a group of multicellular chemosynthetic organisms (similar to that of bacteria living in earth's hot springs and deep sea vents, but relatively more complex, as mushrooms (a fungus with multiple cell fates) and lichens (a symbiosis between a fungus and an alga) are more complex than yeast)? For example, would they have to be sessile, or could they be motile? Vascular or nonvascular? Would their lifestyle necessitate a unique anatomy or mode of reproduction, or might they appear to be just another kind of plant or fungus?

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Multicellular chemotrophs?

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