What conditions would lead a hyperparasite (whose host is itself a parasite) to develop civilization?
A hyperparasite is "a parasite whose host is itself a parasite" and is the closest analogy to an apex predator in the ecology of parasites. Human sociability, intelligence and status as apex predators allowed us to develop civilization. What conditions would pressure a hyperparasite (or hyperparasitoid) to develop social structure, swarm intelligence and eventually civilization?
EDIT: This is not about parasitic aliens that invade Earth to take over humans a la The Puppet Masters, Stargate SG-1, Star Trek, Animorphs, and so forth. Those sorts of parasites would themselves be the hosts of the hyperparasites.
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