How could a collective consciousness evolve?

We have talked about hive minds a lot here and relatively recently, a new idea of hive mind has become popular; the collective consciousness. The basic run down is that a collective consciousness species replaces cells, tissues and organs with ants, specialized ants and groups of specialized ants (or really any small colony animal; termites, bees, wasps, etc). The problem I have hear is figuring out a evolutionary reason why a whole colony would begin to become sentient instead of just remaining as a group.

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How could a collective consciousness evolve?

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