How to justify having overly large irises?

In many famous alien species you see that the iris is so large that it almost appears that it replaces the sclera. What evolutionary history would support the irises growing to that size. The only two requirements of the species are that they need to be humanoid and evolved in jungles.

For the record I am looking for answers other than the typical, "Obviously sexual selection is the answer."

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How to justify having overly large irises?

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