What natural defenses might an ecosystem evolve against dragons?
In a world in which dragons (as animals, minus magical associations) existed, what natural defense mechanisms might other animals or plants evolve in response? How would a creature as devastating like that affect their environment, and what could the rest of the biome do to keep from completely collapsing?
I'm especially interested in ways that other creatures could not only defend themselves from giant flying, fire-breathing reptiles, but also make use of and even exist symbiotically with them, in the same way that flowers use bees to pollenate, etc. Bonus points if these adaptations evolved alongside the development of dragons, rather than in response to the fully-formed appearance of them.
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