Scientific Speculation - Horses went extinct, people domesticated something else.

Assuming that horses went extinct early in history, what would replace the horse, and how would the change in domesticated animal affect the development of the first civilizations?

Note that this question will be followed up with another one based around extrapolating further human development from the situation in the accepted answer.

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Horses went extinct, people domesticated something else. - What is it and what would change?

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